It has an historic church of St. Andrew and St. Mary the Virgin dating from the twelfth century. Simon de Montfort prayed there before the Battle of Lewes. Historian Edward Gibbon (1737–1794) is interred within the church, having died in Fletching while staying with his great friend, John Baker-Holroyd, 1st Earl of Sheffield. In medieval times Fletching was a major producer of bows and arrows, many of which were used at the Battle of Agincourt in 1415.
The school is Fletching CE Primary school. There are two public houses in Fletching: The Griffin Inn (which calls itself a gastropub) and The Rose and Crown. Nearby is The Piltdown Man at Piltdown.
The village was once the home of Jimmy Edwards (1920–1988).