A water beetle is a beetle adapted to living in water. several different types are known, most in or on fresh water. The rest marine species tend to live in the intertidal zone.
Many water beetles carry an air bubble underneath their abdomens, which both provides an air supply, and prevents water from getting into the spiracles. Others have the surface of their exoskeleton modified to form a plastron, or "physical gill", which permits direct gas exchange with the water.
Most families of water beetles have larvae that are also aquatic; many have aquatic larvae and terrestrial adults.
Types that are aquatic in all life stages include whirligig beetles (Gyrinidae), Haliplidae, Noteridae, Amphizoidae, Dytiscidae and Hydroscaphidae
Types in which the adults are not necessarily aquatic include Hydrophilidae, Lutrochidae, Dryopidae, Elmidae, Eulichadidae, Heteroceridae, Limnichidae, Psephenidae, Ptilodactylidae and Sphaeriusidae.