See biography by J. H. Adamson and H. F. Folland (1973).
Inglewood is the eldest son of former Conservative Member of Parliament William Fletcher-Vane and his wife Mary née Proby. He was educated at Eton and at Trinity College, Cambridge, and was called to the bar at Lincoln's Inn in 1975. He married Cressida Pemberton-Pigott in 1986. They have one son, Henry William Frederick Fletcher-Vane born in 1990, and two daughters.
At the European Parliament election, 1984 he stood unsuccessfully in the Durham constituency, but at 1989 European election he was elected as Member of the European Parliament (MEP) for Cumbria and Lancashire North. He lost his seat the 1994 election, but at the European Parliament election, 1999 he was elected for the new North West England constitituency. He did not contest the 2004 election.
He inherited his title from his father in 1989, the same year he was first elected to the European Parliament. He remained a member of the House of Lords after House of Lords Act 1999 after his election by fellow hereditary peers. He was a government whip in the Lords from 1994-95, serving as Deputy Chief Whip in from January to July 1995.. He was then appointed as a junior minister in the Department of National Heritage, serving until the Conservatives lost office at the 1997 general election. He has been a Deputy Lieutenant of Cumbria since 1993.
In 2006 Inglewood suggested eating grey squirrels as a method of preserving the native red squirrel.