Scales have been modified over time to serve other functions such as 'eyelash' fringes, and protective covers for the eyes with the most distinctive modification being the rattle of the North American rattlesnakes.
Snakes periodically moult their scaly skins and acquire new ones. This permits replacement of old worn out skin, disposal of parasites and is thought to allow the snake to grow. The arrangement of scales is used to identify snake species.
Snakes have been part and parcel of culture and religion. Vivid scale patterns have been thought to have influenced early art. The use of snake-skin in manufacture of purses, apparel and other articles led to large-scale killing of snakes, giving rise to advocacy for use of artificial snake-skin. Snake scales are also to be found as motifs in fiction, video games and films.
Snake skin and scales help retain moisture in the animal's body.
Snake scales are formed by the differentiation of the snake's underlying skin or epidermis. Each scale has an outer surface and an inner surface. The skin from the inner surface hinges back and forms a free area which overlaps the base of the next scale which emerges below this scale.
A snake hatches with a fixed number of scales. The scales do not increase in number as the snake matures nor do they reduce in number over time. The scales however grow larger in size and may change shape with each moult.
Snakes have smaller scales around the mouth and sides of the body which allow expansion so that a snake can consume prey of much larger width than itself.
Snake scales are made of keratin, the same material that hair and fingernails are made of. They are cool and dry to touch.
Certain primitive snakes such as boas, pythons and certain advanced snakes such as vipers have small scales arranged irregularly on the head. Other more advanced snakes have special large symmetrical scales on the head called shields or plates.
Snake scales occur in variety of shapes. They may be :
Another example of differentiation of snake scales is a transparent scale called the brille or spectacle which covers the eye of the snake. The brille is often referred to as a fused eyelid. It is shed as part of the old skin during moulting.
The most distinctive modification of the snake scale is the rattle of rattlesnakes, such as those of the genera Crotalus and Sistrurus. The rattle is made up of a series of loosely linked, interlocking chambers that when shaken, vibrate against one another to create the warning signal of a rattlesnake. Only the bottom button is firmly attached to the tip of the tail.
At birth, a rattlesnake hatchling has only a small button or 'primordial rattle' which is firmly attached to the tip of the tail. The first segment is added when the hatchling sheds its skin for the first time. A new section is added each time the skin is shed until a rattle is formed. The rattle grows as the snake ages but segments are also prone to breaking off and hence the length of a rattle is not a reliable indicator of the age of a snake.
Some snakes have the ability to change the hue of their scales slowly. This is typically seen in cases where the snake becomes lighter or darker with change in season. In some cases, this change may take place between day and night.
The shedding of scales is called ecdysis, or, in normal usage moulting or sloughing. In the case of snakes, the complete outer layer of skin is shed in one layer. Snake scales are not discrete but extensions of the epidermis hence they are not shed separately, but are ejected as a complete contiguous outer layer of skin during each moult, akin to a sock being turned inside out.
Moulting serves a number of functions - firstly, the old and worn skin is replaced, secondly, it helps get rid of parasites such as mites and ticks. Renewal of the skin by moulting is supposed to allow growth in some animals such as insects, however this view has been disputed in the case of snakes.
Moulting is repeated periodically throughout a snake's life. Before a moult, the snake stops eating and often hides or moves to a safe place. Just before shedding, the skin becomes dull and dry looking and the eyes become cloudy or blue-colored. The inner surface of the old outer skin liquefies. This causes the old outer skin to separate from the new inner skin. After a few days, the eyes clear and the snake "crawls" out of its old skin. The old skin breaks near the mouth and the snake wriggles out aided by rubbing against rough surfaces. In many cases the cast skin peels backward over the body from head to tail, in one piece like an old sock. A new, larger, and brighter layer of skin has formed underneath.
An older snake may shed its skin only once or twice a year, but a younger, still-growing snake, may shed up to four times a year. The discarded skin gives a perfect imprint of the scale pattern and it is usually possible to identify the snake if this discard is reasonably complete and intact.
The dorsal (or body) scales on the snake's body are arranged in rows along the length of their bodies. Adjacent rows are diagonally offset from each other. Most snakes have an odd number of rows across the body though certain species have an even number of rows e.g. Zaocys spp. In the case of some aquatic and marine snakes, the scales are granular and the rows cannot be counted.
The number of rows range from ten in Tiger Ratsnake Spilotes pullatus; thirteen in Dryocalamus, Liopeltis, Calamaria and Asian coral snakes of genus Calliophis; 65 to 75 in pythons; 74 to 93 in Kolpophis and 130 to 150 in Acrochordus. The majority of the largest family of snakes, the Colubridae have 15, 17 or 19 rows of scales.
The maximum number of rows are in mid-body and they reduce in count towards the head and on the tail.
Identification of scales is most conveniently begun with reference to the nostril which is easily identified on the snake. There are two scales enclosing the nostril which are called the nasals. The outer nasal (near the snout) is called the prenasal while the inner nasal (near the eye) is called the postnasal. Along the top of the snout connecting the nasals on both sides of the head are scales called internasals.
Between the two prenasals is a scale at the tip of the snout called the rostral scale.
The scales around the eye are called circumorbital scales and are named as 'ocular' scales but with appropriate prefix. The ocular scale proper is a transparent scale covering the eye which is called the spectacle, brille or eyecap..
The circumorbital scales towards the snout or the front are called preocular scales, those towards the rear are called postocular scales and those towards the upper or dorsal side are called as supraocular scales. Circumorbital scales towards the ventral or lower side, if any, are called as subocular scales.
Between the preocular and the postnasal scales are the loreal scales.
The scales along the lips of the snake are called as labials. Those on the upper lip are called supralabials while those on the lower labial are called infralabials.
Between the eyeballs on top of the head, adjacent to the supraoculars are the frontal scales. The prefrontal scales are the scales connected to the frontals towards the tip of the snout which are in contact with the internasals.
The back of the top of the head has scales connected to the frontal scales called as the parietal scales. At the sides of the back of the head are scales called temporal scales.
On the underside of the head, a snake has an anterior scale called as the mental scale. Connected to the mental scales and all along the lower jaws are the infralabials. Along the lower jaw connected to infralabials are a pair of shields called the anterior chin shields. Next to the anterior chin shields, further back along the jaw are another pair of shields called the posterior chin shields.
The mental groove is a longitudinal groove on the underside of the head between large, paired chin shields and smaller gular scales.
The scales on the body of the snake are called the dorsal or costal scales. Sometimes there is a special row of large scales along the top of the back of the snake, i.e, the uppermost row, called the vertebral scales.
The enlarged scales on the belly of the snake are called ventral scales or gastrosteges.
In "advanced" (Caenophidian) snakes, the broad belly scales and rows of dorsal scales correspond to the vertebrae, allowing scientists to count the vertebrae without dissection.
At the end of the ventral scales of the snake is an anal plate which protects the opening to the cloaca (a shared opening for waste and reproductive material to pass) on the underside near the tail. This anal scale may be single or divided into a pair. The part of the body beyond the anal scale is considered to be the tail.
Sometimes snakes have enlarged scales, either single or paired, under the tail; these are called subcaudals or urosteges. These subcaudals may be smooth or keeled as in Bitis arietans somalica. The end of the tail may simply taper into a tip (as in the case of most snakes), it may form a spine (as in Acanthophis), end in a bony spur (as in Lachesis), a rattle (as in Crotalus), or a rudder as seen in many sea snakes.
Sources. Details for this section have been sourced from scale diagrams in Malcolm Smith. Details of scales of Buff-striped Keelback have been taken from Daniels.
In certain areas in North America, where the diversity of snakes is not too large, easy keys based on simple identification of scales have been devised for the lay public to distinguish poisonous snakes from non-poisonous snakes. In other places with large biodiversity, such as Myanmar, publications caution that venomous and non-venomous snakes cannot be easily distinguished apart without careful examination.
Identification of snakes by reference to scales seems an arcane art to the amateur naturalist because the nomenclature of snake scales seems esoteric and, more importantly, the snakes need to be caught and the head and body examined closely in hand for identification. The advent of high definition digital cameras means that images taken of snakes, if of appropriate definition and of the correct position, would show scales that can be examined, distinguished and counted without the need for catching and handling.
The scales patterning may also be used for individual identification in field studies. Clipping of specific scales to mark individual snakes is a popular approach to population estimation by mark and recapture techniques.
A point to note is that such identification requires a fair degree of knowledge about snakes, their taxonomy, snake-scale nomenclature as well as familiarity and access to authoritative scientific texts on snakes.
In certain regions, experts use presence or absence of certain scales to distinguish between non-venomous and venomous scales, with well-understood exceptions. For example, in relation to venomous snakes of Myanmar, the distribution of snakes permits the use of the presence or absence of loreal scales to distinguish between relatively harmless Colubrids and lethally venomous Elapids.
Please note that this rule-of-hand applies to terrestrial venomous snakes only and that vipers are an exception since they cannot be so classified due to the large number of small scales on the head. Subsequently a further check is to be carried out in the case of identified Colubrids to exclude known poisonous members of their family such as Rhabdophis species.
In South Asia, it is advisable to kill the snake which has bitten a person and carry it along to the hospital for possible identification by medical staff using scale diagrams so that an informed decision can be taken them as to whether and which anti-venin is to be administered.
Snakeskin, with its highly periodic cross-hatch or grid patterns, appeals to people's aesthetics and have been used to manufacture many leather articles including fashionable accessories. The use of snakeskin has however endangered snake populations and resulted in international restrictions in trade of certain snake species and populations in the form of CITES provisions. Animal lovers in many countries now propagate the use of artificial snakeskin instead, which are easily produced from embossed leather, patterned fabric, plastics and other materials.
Snake scales occur as a motif regularly in computer action games.
A snake scale was portrayed as a clue in the 1982 film Blade Runner.