Snakes constitute the suborder Serpentes (or Ophidia). In most snakes limbs are entirely lacking, but a few have traces of hind limbs. The skin, which is covered with horny scales, is shed, usually several times a year. The extremely long, narrow body is associated with distinctive internal features. The number of vertebrae is much larger than in most vertebrates, paired internal organs are arranged linearly rather than side by side, and only one lung is developed, except in members of the boa family, which have two lungs. The jaws of snakes are loosely jointed and extremely flexible. The pointed, backward-curved teeth are fused to the supporting bones of the head. There are no ears or movable eyelids. Snakes have good vision. They do not hear airborne sound waves, but can perceive low-frequency vibrations (100-700 Hz) transmitted from the ground to the bones of the skull. A chemosensory organ opens into the roof of the mouth; it receives stimuli from the forked tongue that constantly tastes the surroundings as the animal moves along. Snakes have no larynx or vocal chords, but are capable of producing a hissing sound.
Locomotion and LimblessnessA snake moves by means of muscular contraction, which can produce several types of locomotion, the commonest types being undulation and straight-line movement. Straight-line movement is aided by the ventral plates, elongated scales on the abdomen that overlap with their open ends pointing toward the tail. These plates can be moved forward by means of muscles attached to the ribs.
It is believed that snakes are descended from lizards, but how and why they evolved toward limblessness is uncertain. Some paleontologists have held that limblessness was an evolutionary advantage in the dense vegetation that formed the early environment of snakes, or that it developed to facilitate burrowing habits, but others believe that the earliest snakes evolved in a marine environment and are descended from marine lizards. Support for the latter view comes from recent discovery of the most primitive snake now known, a fossil specimen with two short but well-developed hind legs found in marine sediments in Israel. It lived in a shallow sea 95 million years ago.
PredationSmall snakes feed on insects and larger ones on proportionately larger animals. Their teeth are designed for catching and holding prey, but not for chewing. The construction of the jaws, the ribs, and the expandable skin enable them to swallow very large prey whole. Some snakes capture animals by pinning them to the ground; some—the constrictors—crush them by wrapping their bodies around them and squeezing; still others—the venomous snakes—inject poison into their victims. The poison, or venom, is produced by modified salivary glands from which it passes through either a groove or a hollow bore in the fangs, the enlarged, specialized teeth found in venomous snakes. A snake may bite a person when threatened or alarmed; if the snake is venomous the bite can sometimes prove fatal (see snakebite). Only by familiarity with the appearance of particular species, or by examination of the fangs, can the venomous snakes be distinguished from the harmless ones.
ReproductionFertilization is internal in snakes; as in lizards, the males have paired copulatory organs, either of which may be used in mating. Females of some species can store sperm for several years to insure future fertilization. In most species the female lays eggs; in some the eggs are incubated and hatched within the mother's body; in a few there is true viviparity, or live birth, with the young nourished by means of a placenta rather than an egg. Some egg-laying snakes brood the eggs, but there is no parental care of the young.
The approximately 2,700 snake species, of which about four fifths are nonvenomous, are distributed throughout the temperate and tropical zones of the world (except in New Zealand, Ireland, and some isolated oceanic islands) and are found in greatest profusion in the tropics. About two thirds of all snake species belong to the family Colubridae; most of these are nonvenomous. Among the harmless colubrid snakes of North America are the garter snakes (including the ribbon snake), the water snakes, the green, or grass, snakes, the black snakes, the racers, the king snakes (including the milk snake), and the bull, hognose, and rat snakes. The family Boidae (boas and pythons) includes the world's largest snakes, the South American anaconda and the Asian reticulated python, as well as the smaller boa constrictor and the tree and sand boas.
Most poisonous New World snakes belong to the pit viper family; these include the copperhead, water moccasin, rattlesnake, fer-de-lance, and bushmaster. Venomous Old World snakes are the true vipers, including the adder and the asp, and members of the cobra family, including the mamba of Africa and the krait of Asia. The poisonous coral snakes of the New World also belong to this family. The venomous sea snakes inhabit tropical oceans.
Snakes are of major importance as pest controllers because of their extensive predation on destructive mammals such as rats and mice. Some, like the sea snakes and pythons, are highly regarded as food in Asia but, although most are probably edible, snakes are not widely used for meat. The skin is often used for belts, bags, and shoes. Venom is removed from snakes for use in treating certain diseases and to make antivenin for snakebites.
See also snake worship.
Snakes are classified in the phylum Chordata, subphylum Vertebrata, class Reptilia, order Squamata, suborder Serpentes.
See A. H. and A. A. Wright, A Handbook of Snakes of the United States and Canada (2 vol., 1957); K. L. Williams and V. Wallach, Snakes of the World (2 vol., 1990).
The Snake was explored by the Lewis and Clark expedition (1803-6) and was of major importance in U.S. expansion into the Pacific Northwest. The river is a major source of electricity, having numerous hydroelectric power plants. The upper and middle courses of the Snake and its tributaries are much used for irrigation by private projects (one of the most notable being at Twin Falls) and by projects of the U.S. Bureau of Reclamation, including the Minidoka project, the Boise project, the Palisades project, and the Owyhee project. Four navigation and hydroelectric power projects along the lower Snake provide slack water navigation from the mouth of the Snake 140 mi (225 km) upstream to Lewiston, Idaho. The projects are linked with the navigation system on the Columbia River. The late 1990s brought efforts to restore portions of the river by removing gravel and establishing new islands.
Any of about 200 snake species of about 38 genera, especially Natrix and Nerodia, classified within the family Colubridae, found worldwide except in South America. Most species have a stout body with dark blotches or streaks and ridged scales. Some are similar in appearance to venomous species. They kill fishes and amphibians with a nonvenomous bite. The New World species live in or near water and bear live young; European species are less water-dependent and lay eggs. In defense they inflate the head, strike, and release a foul secretion. Average length is about 3 ft (1 m); some Old World species reach 6 ft (1.8 m).
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Region, south and southwestern U.S. It is characterized by a warm climate, rapid population growth since 1970, and relatively conservative voting patterns. Comprising 15 states, it extends from Virginia and Florida in the southeast through Nevada in the southwest, and includes southern California.
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Internal and external features of a snake.
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Any of some 50 species (family Hydrophiidae) of venomous, marine snakes with an oarlike tail and flattened body. Most are found along coasts and in estuaries of Australia and Asia, sometimes basking on the surface in a large group, though the yellow-bellied, or pelagic, sea snake ranges throughout the Pacific. The nostrils, usually on top of the snout, have valvelike closings. The body of several species is much thicker than the head and neck. Most species are 3–4 ft (1–1.2 m) long; Laticauda semifasciata, a Japanese delicacy, may be twice as long. Though generally slow to strike, their venom may be lethal.
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Aesculapian snake (Elaphe longissima).
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Common king snake (Lampropeltis getula).
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Indigo snake (Drymarchon corais)
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Any of three or four species (genus Heterodon, family Colubridae) of harmless North American snakes named for their upturned snout, which is used for digging. When threatened, they flatten the head and neck, then strike with a loud hiss, but rarely bite. If their bluff fails, they roll over, writhing, and then act dead, with mouth open and tongue lolling. They eat chiefly toads and frogs. Heavy-bodied and blotchy, they are usually about 24 in. (60 cm) long. Though not adders, they are sometimes called puff adders or blow snakes.
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Garter snake (Thamnophis).
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Any of about 90 species of strongly patterned burrowing elapids. “True” forms are limited to the New World, chiefly the tropics, but similar species live in Asia and Africa. Secretive and docile, coral snakes rarely bite when handled, but the venom of some can kill a person. Most prey on other snakes. More than 50 species in the largest genus, Micrurus, range from the southern U.S. to Argentina. They are ringed with red, black, and yellow or white. The eastern coral snake, or harlequin snake (Micrurus fulvius), ranges from North Carolina and Missouri in the U.S. to northeastern Mexico and is about 30 in. (76 cm) long, with wide bands of red and black separated by yellow. The rhyme “Red on yellow, dangerous fellow” distinguishes it from similarly coloured but harmless species.
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Any of several species of all-black or nearly all-black snakes. Australian black snakes are in the elapid genus Pseudeschis. The black snake of Australian wetlands (P. porphyriacus) grows to an average length of 5 ft (1.5 m). If annoyed, it expands its neck, cobra fashion. Its venom is rarely fatal. Other Australian black snakes are the mulga snake (P. australis) and the spotted black snake (P. guttatus). North American black snakes include two species in the family Colubridae: the black racer and the pilot black snake (Elaphe obsoleta).
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Group of closely related North American Indian peoples living in the Great Basin region of the U.S. Their language belongs to the Numic group of the Uto-Aztecan family. The Shoshone are usually divided into four groups: Western (unmounted) Shoshone, centred in eastern Nevada; Northern (mounted) Shoshone of northwestern Utah and southern Idaho; Wind River Shoshone in western Wyoming; and the Comanche, a relatively recent division related to the Wind River peoples, in western Texas. The Western Shoshone traditionally subsisted through hunting and gathering. The Northern Shoshone and Wind River Shoshone probably acquired horses by 1680 and adopted much of Plains Indian culture; they hunted buffalo, used tepees and skin clothing, and warred with other tribes. The Shoshone are closely related to the Ute, Paiute, Gosiute, and Bannock. Early 21st-century population estimates indicated some 19,000 individuals of Comanche descent and an additional 22,000 individuals of Western, Northern, or Wind River Shoshone descent.
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