In early Western physiological theory, one of the four body fluids thought to determine a person's temperament and features. As hypothesized by Galen, the four cardinal humours were blood, phlegm, choler (yellow bile), and melancholy (black bile). The variant mixture of these humours in each person determined his “complexion” or temperament and his mental and physical qualities. The ideal person had the perfectly proportioned mixture of the four fluids; a disproportionate amount of one humour created a personality dominated by one set of related emotions (e.g., a choleric man was easily angered, proud, ambitious, and vengeful).
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Humour marked by the use of morbid, ironic, or grotesquely comic episodes that ridicule human folly. The term came into common use in the 1960s to describe the work of novelists such as Joseph Heller, whose Catch-22 (1961) is an outstanding example; Kurt Vonnegut, particularly in Slaughterhouse Five (1969); and Thomas Pynchon, in V (1963) and Gravity's Rainbow (1973). A film exemplar is Stanley Kubrick's Dr. Strangelove (1963). The term black comedy has been applied to some playwrights in the Theatre of the Absurd, especially Eugène Ionesco.
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Mechanism by which information is received about one's external or internal environment. Stimuli received by nerves, in some cases through specialized organs with receptor cells sensitive to one type of stimulus, are converted into impulses that travel to specialized areas of the brain, where they are analyzed. In addition to the “five senses”—sight, hearing, smell, taste, and touch—humans have senses of motion (kinesthetic sense), heat, cold, pressure, pain, and balance. Temperature, pressure, and pain are cutaneous (skin) senses; different points on the skin are particularly sensitive to each. Seealso chemoreception, ear, eye, inner ear, mechanoreception, nose, photoreception, proprioception, taste, thermoreception, tongue.
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Senses are the physiological methods of perception. The senses and their operation, classification, and theory are overlapping topics studied by a variety of fields, most notably neuroscience, cognitive psychology (or cognitive science), and philosophy of perception. The nervous system has a specific sensory system, or organ, dedicated to each sense.
One commonly recognized catagorisation for human senses is as follows: chemoreception; photoreception; mechanoreception; and thermoception. Indeed, all human senses fit into one of these four categories.
Different senses also exist in other organisms, for example electroreception.
A broadly acceptable definition of a sense would be "a system that consists of a group sensory cell types that responds to a specific physical phenomenon, and that corresponds to a particular group of regions within the brain where the signals are received and interpreted." Disputes about the number of senses arise typically regarding the classification of the various cell types and their mapping to regions of the brain.
Sight or vision is the ability of the brain and eye to detect electromagnetic waves within the visible range (light) interpreting the image as "sight." There is disagreement as to whether this constitutes one, two or three senses. Neuroanatomists generally regard it as two senses, given that different receptors are responsible for the perception of colour (the frequency of photons of light) and brightness (amplitude/intensity - number of photons of light). Some argue that stereopsis, the perception of depth, also constitutes a sense, but it is generally regarded as a cognitive (that is, post-sensory) function of brain to interpret sensory input and to derive new information. The inability to see is called blindness.
Hearing or audition is the sense of sound perception. Since sound is vibrations propagating through a medium such as air, the detection of these vibrations, that is the sense of the hearing, is a mechanical sense akin to a sense of touch, albeit a very specialized one. In humans, this perception is executed by tiny hair fibres in the inner ear which detect the motion of a membrane which vibrates in response to changes in the pressure exerted by atmospheric particles within a range of 20 to 22000 Hz, with substantial variation between individuals. Sound can also be detected as vibrations conducted through the body by tactition. Lower and higher frequencies than that can be heard are detected this way only. The inability to hear is called deafness.
The four well-known receptors detect sweet, salt, sour, and bitter, although the receptors for sweet and bitter have not been conclusively identified. A fifth receptor, for a sensation called umami, was first theorised in 1908 and its existence confirmed in 2000. The umami receptor detects the amino acid glutamate, a flavour commonly found in meat and in artificial flavourings such as monosodium glutamate.
Smell or olfaction is the other "chemical" sense. Unlike taste, there are hundreds of olfactory receptors, each binding to a particular molecular feature. Odour molecules possess a variety of features and thus excite specific receptors more or less strongly. This combination of excitatory signals from different receptors makes up what we perceive as the molecule's smell. In the brain, olfaction is processed by the olfactory system. Olfactory receptor neurons in the nose differ from most other neurons in that they die and regenerate on a regular basis. The inability to smell is called anosmia.
Touch, also called tactition, mechanoreception or somatic sensation, is the sense of pressure perception, generally in the skin. There are a variety of nerve endings that respond to variations in pressure (e.g., firm, brushing, and sustained). The inability to feel anything or almost anything is called anesthesia. Paresthesia is a sensation of tingling, pricking, or numbness of a person's skin with no apparent long term physical effect.
Balance, Equilibrioception, or vestibular sense, is the sense which allows an organism to sense body movement, direction, and acceleration, and to attain and maintain postural equilibrium and balance. The organ of equilibrioception is the vestibular labyrinthine system found in both of the inner ears. Technically this organ is responsible for two senses, angular momentum and linear acceleration (which also senses gravity), but they are known together as equilibrioception.
The vestibular nerve conducts information from the three semicircular canals, corresponding to the three spatial planes, the utricle, and the saccule. The ampulla, or base, portion of the three semicircular canals each contain a structure called a crista. These bend in response to angular momentum or spinning. The saccule and utricle, also called the "otolith organs", sense linear acceleration and thus gravity. Otoliths are small crystals of calcium carbonate that provide the inertia needed to detect changes in acceleration or gravity.
Thermoception is the sense of heat and the absence of heat (cold) by the skin and including internal skin passages. The thermoceptors in the skin are quite different from the homeostatic thermoceptors in the brain (hypothalamus) which provide feedback on internal body temperature.
Proprioception, the kinesthetic sense, provides the parietal cortex of the brain with information on the relative positions of the parts of the body. Neurologists test this sense by telling patients to close their eyes and touch the tip of a finger to their nose. Assuming proper proprioceptive function, at no time will the person lose awareness of where the hand actually is, even though it is not being detected by any of the other senses. Proprioception and touch are related in subtle ways, and their impairment results in surprising and deep deficits in perception and action.
Nociception (physiological pain) signals near-damage or damage to tissue. The three types of pain receptors are cutaneous (skin), somatic (joints and bones) and visceral (body organs). It was believed that pain was simply the overloading of pressure receptors, but research in the first half of the 20th century indicated that pain is a distinct phenomenon that intertwines with all of the other senses, including touch. Pain was once considered an entirely subjective experience, but recent studies show that pain is registered in the anterior cingulate gyrus of the brain.
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