The origins of the PUP suite lie in two developments; in the same events in the early 1970s as the very earliest stage of the development of TCP/IP (see History of the Internet), and the creation of the Ethernet local area network at PARC. However, the development of PUP split off because Xerox PARC wished to move ahead with implementation, for in-house use. The fundamental design of the PUP suite was substantially complete by 1974.
In the 1980s Xerox used PUP as the base for the Xerox Network Services (XNS) protocol suite; some of the protocols in the XNS suite (e.g. the Internetwork Datagram Protocol) were lightly modified versions of the ones in the PUP suite, but others are quite different, reflecting the experience gained with PUP.
The main internetwork layer protocol was PUP, which roughly corresponds to the Internet Protocol (IP) layer in TCP/IP. A full PUP network address consisted of an 8-bit network number, an 8-bit host number, and a 16-bit socket number. The network number had a particular special value which meant 'this network', for use by hosts which did not (yet) know their network number.
Unlike TCP/IP, socket fields were part of the full network address in the PUP header, so that upper-layer protocols did not need to implement their own demultiplexing; PUP also supplied packet types (again, unlike IP). Also, an optional 2-byte checksum covered the entire packet.
PUP packets were up to 554 bytes long (including the 20 byte PUP header), and the checksum. This was a smaller packet size than IP, which requires all hosts to support at least 576 (but supports packets of up to 65K bytes, if the hosts support them); individual PUP host pairs on a particular network might use larger packets, but no PUP router was required to handle them. Larger packets could be fragmented.
PUP also included a simple echo protocol at the internetwork layer, similar to IP's ping, but operating at a lower level.
To establish a transport connection, two protocols came into play. The first, the Rendezvous and Termination Protocol (RTP), which was used to initiate communication between two entities, as well as manage and terminate the connection. The second was the primary transport layer protocol, Byte Stream Protocol (BSP), which was analogous to TCP.
Once RTP had started the connection, BSP took over and managed the data transfer. Like TCP, BSP's semantics and operation were in terms of bytes; this was discarded in favour of packets for the equivalent protocol in XNS, Sequenced Packet Protocol.
Others were novel, including protocols for printer spooling, copying disk packs, page-level remote access to file servers, name lookup, remote management, etc (although some of these capabilities had been seen before, e.g. the ARPANET already made heavy use of remote management for controlling the Interface Message Processors which made it up).
In showing that internetworking ideas were feasible, in being influential in the early work on TCP/IP, and as the foundation for the later XNS protocols, PUP was very influential. However, its biggest impact was probably as a key component of the office of the future model first demonstrated at Xerox PARC; that demonstration would not have been anything like as powerful as it was without all the capabilities that a working internetwork provided.
The Gateway Information Protocol's descendant, RIP, (somewhat modified to match the syntax of addresses of other protocol suites), remains in wide use today in other protocol suites. One version of RIP served as one of the initial so-called interior gateway protocols for the growing Internet, before the arrival of the more modern OSPF and IS-IS. It is still in use as an interior routing protocol, in small sites with simple requirements.