Zoomusicology is often a subfield of zoosemiotics.
Writers on music semiology include Kofi Agawu (on topical theory, Schenkerian analysis), Robert Hatten (on topic, gesture), Raymond Monelle (on topic, musical meaning), Jean-Jacques Nattiez (on introversive taxonomic analysis and ethnomusicological applications), Anthony Newcomb (on narrativity), and Eero Tarasti (generally considered the founder of musical semiotics).
Roland Barthes, himself a semiotician and skilled amateur pianist, wrote about music in Image-Music-Text, The Responsibilities of Form, and Eiffel Tower, though he did not consider music to be a semiotic system.
Signs, meanings in music, happen essentially through the connotations of sounds, and through the social construction, appropriation and amplification of certain meanings associated with these connotations. The work of Philip Tagg (Ten Little Tunes, Fernando the Flute) provides one of the most complete and systematic analysis of the relation between musical structures and connotations in western and especially popular, television and film music. The work of Leonard Meyer in Style and Music theorizes the relationship between ideologies and musical structures and the phenomenons of style change, and focuses on romanticism as a case study.