Ernst Heinrich Philipp August Haeckel (February 16, 1834 — August 9, 1919), also written von Haeckel, was an eminent German biologist, naturalist, philosopher, physician, professor and artist who discovered, described and named thousands of new species, mapped a genealogical tree relating all life forms, and coined many terms in biology, including phylum, phylogeny, ecology and the kingdom Protista. Haeckel promoted and popularized Charles Darwin's work in Germany and developed the controversial recapitulation theory ("ontogeny recapitulates phylogeny") claiming that an individual organism's biological development, or ontogeny, parallels and summarizes its species' entire evolutionary development, or phylogeny.
The published artwork of Haeckel includes over 100 detailed, multi-colour illustrations of animals and sea creatures (see: Kunstformen der Natur, "Artforms of Nature"). As a philosopher, Ernst Haeckel wrote Die Welträthsel (1895-1899, in English, The Riddle of the Universe, 1901), the genesis for the term "world riddle" (Welträthsel); and Freedom in Science and Teaching to support teaching evolution.
In the United States, Mount Haeckel, a summit in the Eastern Sierra Nevada, overlooking the Evolution Basin, is named in his honor, as are another Mount Haeckel, a summit in New Zealand; and the asteroid 12323 Häckel.
Ernst Haeckel was born on February 16, 1834, in Potsdam (then part of Prussia). In 1852, Haeckel completed studies at Cathedral High School (Domgymnasium) of Merseburg. He then studied medicine in Berlin, particularly with Albert von Kölliker, Franz Leydig, Rudolf Virchow (with whom he later worked briefly as assistant), and with anatomist-physiologist Johannes Peter Müller (1801-1858). In 1857, Haeckel attained a doctorate in medicine (M.D.), and afterwards he received a license to practice medicine. The occupation of physician appeared less worthwhile to Haeckel, after contact with suffering patients.
Haeckel studied under Carl Gegenbaur at the University of Jena for three years, earning a doctorate in zoology, before becoming a professor of comparative anatomy at the University of Jena, where he remained 47 years, from 1862-1909. Between 1859 and 1866, Haeckel worked on many invertebrate groups, including radiolarians, poriferans (sponges) and annelids (segmented worms). During a trip to the Mediterranean, Haeckel named nearly 150 new species of radiolarians.
Haeckel named thousands of new species from 1859 to 1887.
From 1866 to 1867, Haeckel made an extended journey to the Canary Islands and during this period, met with Charles Darwin, in 1866 at Down House in Kent, Thomas Huxley and Charles Lyell. In 1867, he married Agnes Huschke. Their son Walter was born in 1868, their daughters Elizabeth in 1871 and Emma in 1873. In 1869, he traveled as a researcher to Norway, in 1871 to Dalmatia, and in 1873 to Egypt, Turkey, and to Greece. Haeckel retired from teaching in 1909, and in 1910 he withdrew from the Evangelical church. Haeckel's wife, Agnes, died in 1915, and Ernst Haeckel became substantially more frail, with a broken leg (thigh) and broken arm. He sold the mansion Medusa ("Villa Medusa") in 1918 to the Carl Zeiss foundation. Ernst Haeckel died on August 9, 1919.
There is no doubt that the course and character of the feared "European War"...will become the first world war in the full sense of the word.
The "European War" became known as "The Great War", and it was not until 1931, with the beginning realization that another global war might be possible, that there is any other recorded use of the term "First World War".
Haeckel was a zoologist, an accomplished artist and illustrator, and later a professor of comparative anatomy. Although Haeckel's ideas are important to the history of evolutionary theory, and he was a competent invertebrate anatomist most famous for his work on radiolaria, many speculative concepts that he championed are now considered incorrect. For example, Haeckel described and named hypothetical ancestral microorganisms that have never been found.
He was one of the first to consider psychology as a branch of physiology. He also proposed many now ubiquitous terms including "phylum", "phylogeny", "ecology" ("oekologie"), and proposed the kingdom Protista in 1866. His chief interests lay in evolution and life development processes in general, including development of nonrandom form, which culminated in the beautifully illustrated Kunstformen der Natur (Art forms of nature). Haeckel did not support natural selection, rather believing in a Lamarckian inheritance of acquired characteristics (Lamarckism).
Haeckel advanced the "recapitulation theory" which proposed a link between ontogeny (development of form) and phylogeny (evolutionary descent), summed up in the phrase "ontogeny recapitulates phylogeny". His concept of recapitulation has been disputed in the form he gave it (now called "strong recapitulation"). "Strong" recapitulation hypothesis views ontogeny as repeating forms of the ancestors, while "weak" recapitulation means that what is repeated (and built upon) is the ancestral embryonic development process. He supported the theory with embryo drawings that have since been shown to be oversimplified and in part inaccurate, and the theory is now considered an oversimplification of quite complicated relationships. Haeckel introduced the concept of "heterochrony", which is the change in timing of embryonic development over the course of evolution.
Haeckel was a flamboyant figure. He sometimes took great (and non-scientific) leaps from available evidence. For example, at the time that Darwin first published On the Origin of Species by Means of Natural Selection (1859), no remains of human ancestors had yet been found. Haeckel postulated that evidence of human evolution would be found in the Dutch East Indies (now Indonesia), and described these theoretical remains in great detail. He even named the as-of-yet unfound species, Pithecanthropus alalus, and charged his students to go find it. (Richard and Oskar Hertwig were two of Haeckel's many important students.)
One student did find the remains: a young Dutchman named Eugene Dubois went to the East Indies and dug up the remains of Java Man, the first human ancestral remains ever found. These remains originally carried Haeckel's Pithecanthropus label, though they were later reclassified as Homo erectus.
Some creationists have claimed that Darwin relied on Haeckel's embryo drawings as proof of evolution
to support their argument that Darwin's theory is therefore illegitimate and possibly fraudulent. This claim ignores the fact that Darwin published On the Origin of Species in 1859, and The Descent of Man in 1871, whereas Haeckel's famous embryo drawings did not appear until 1874 (8 species). In The Descent of Man Darwin used only two embryo drawings, neither taken from Haeckel.
It has been claimed that Ernst Haeckel sent a letter to the January 9 1909 publication of Münchener Allgemeine Zeitung (translated: Munich General Newspaper) which reads, as translated: "A small portion of my embryo-pictures (possibly 6 or 8 in a hundred) are really (in Dr Brass’s sense of the word) 'falsified' — all those, namely, in which the disclosed material for inspection is so incomplete or insufficient that one is compelled in a restoration of a connected development series to fill up the gaps through hypotheses, and to reconstruct the missing members through comparative syntheses. What difficulties this task encounters, and how easily the drafts--man may blunder in it, the embryologist alone can judge."
Darwin’s 1859 book On the Origin of Species had immense popular influence, but although its sales exceeded its publisher's hopes it was a technical book rather than a work of popular science: long, difficult and with few illustrations. One of Haeckel's books did a great deal to explain his version of "Darwinism" to the world. It was a bestselling, provocatively illustrated book in German, titled Natürliche Schöpfungsgeschichte, published in Berlin in 1868, and translated into English as The History of Creation in 1876. It was frequently reprinted until 1926.
Haeckel argued that human evolution consisted of precisely 22 phases, the 21st — the "missing link" — being a halfway step between apes and humans. He even formally named this missing link Pithecanthropus alalus, translated as "ape man without speech." (The missing link was what the Dutchman Eugène Dubois, discoverer of Homo erectus, would later resolve to find.)
Haeckel's entire literary output was extensive, working as a professor at the University of Jena for 47 years, and even at the time of the celebration of his 60th birthday at Jena in 1894, Haeckel had produced 42 works with nearly 13,000 pages, besides numerous scientific memoirs and illustrations.
Haeckel's monographs include:
As well as several Challenger reports:
Among his many books, Ernst Haeckel wrote:
Books of travel: