beaver, either of two large aquatic rodents, Castor fiber and Castor canadensis, known for their engineering feats. They were once widespread in N and central Eurasia except E Siberia, and in North America from the arctic tree line to the S United States. The mountain beaver of W North America is not a true beaver, but a nonaquatic rodent of a different family.

The beaver is the largest living rodent except the capybara, and is distinguished by its extremely broad, horizontally flattened tail. Beavers are 3 to 4 ft (91-120 cm) long, including the tail (12 in./30.5 cm long, 6 in./15.2 cm wide), and about 15 in. (38 cm) high at the shoulder; they usually weigh about 60 lb (27 kg). Their long, dense fur is reddish brown to nearly black; the naked, scaly tail is black. Both sexes have scent glands located in a pouch in the anal region. The musky secretion, castoreum, which may function as a sexual attractant, was once believed to have medicinal properties, and the glands, or castors, were of commercial value. Beavers have been extensively trapped for their pelts, once considered the most valuable of furs, and were exterminated over a large part of their range. Because of their great importance in maintaining the natural environment, they have been reintroduced in many areas of North America and Russia and are now increasing in numbers.

Beavers build lodges up to 3 ft (91 cm) high and 5 ft (1.5 m) wide of sticks and mud; the entrances are below water level, with ramps leading to the living quarters, located on a platform above water level. They may also build burrows in banks with underwater entrances. They create deep ponds, or maintain the water level in old ones, by building dams across streams. These are made of sticks and logs, and the upper surfaces are reinforced with stones and mud. Materials are gathered by collecting wood and felling small trees by gnawing; often the beavers dig canals for floating these to the right spot. Most, if not all, of these activities are done mechanically, as a result of instinct; captive animals persist in building useless dams, and even in the wild beavers will attempt to reinforce solid, manmade dams with sticks.

Although they form monogamous families and live in colonies, there is little social contact among beavers and they work independently. A colony consists of a cluster of lodges, each occupied by a family of the parents and their last two litters. The beavers sleep by day and spend the night foraging for food and building or repairing their structures. They feed on a variety of aquatic and shore plants, surviving in winter largely on bark. Sticks for winter food are stored in the lodges and underwater. Excellent swimmers, they can stay underwater for up to fifteen minutes. When alarmed, a beaver slaps the water with its tail, making a loud noise that sends other beavers hurrying to the safety of deep water. Females give birth to two to eight young in the spring; these mature in two years. Beavers are responsible for creating many of the woodland ponds that support lush vegetation and eventually become meadows.

Classification

Beavers are classified in the phylum Chordata, subphylum Vertebrata, class Mammalia, order Rodentia, family Castoridae.

Either species of the aquatic rodent family Castoridae (genus Castor), both of which are well known for building dams. Beavers are heavyset and have short legs and large, webbed hind feet. They grow as large as 4 ft (1.3 m) long, including the 1-ft (30-cm) tail, and as heavy as 66 lb (30 kg). Beavers build their dams of sticks, stones, and mud in small rivers, streams, and lakes, often producing sizable ponds. With their powerful jaws and large teeth, they can fell medium-size trees, whose branches they use in their dams and whose tender bark and buds they eat. One or more family groups share a dome-shaped stick-and-mud lodge built in the water, with tunnel entrances below water level. American beavers (C. canadensis) range from northern Mexico to the Arctic. Their prized pelts stimulated the exploration of western North America, and by 1900 beavers were trapped to near extinction. Eurasian beavers (C. fiber) are now found in only a few locations, including the Elbe and Rhône drainages of Europe. The mountain beaver of the Pacific Northwest is unrelated.

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Beavers are two primarily nocturnal, semi-aquatic species of rodent, one native to North America and one to Europe. They are known for building dams, canals, and lodges (homes). They are the second-largest rodent in the world (after the capybara). Their colonies create one or more dams to provide still, deep water to protect against predators, and to float food and building material. The North American population was 60-400 million, but as of 1988, 6-12 million, due to being hunted for fur, for their glands used as medicine and perfume, and because their harvesting of trees and flooding of waterways may interfere with other land uses.

General

Beavers are known for their natural trait of building dams in rivers and streams, and building their homes (known as beaver lodges) in the resulting pond. Beavers also build canals to float build materials that are difficult to haul over land. They use powerful front teeth to cut trees and plants that they use for building and for food.

They are known for their "danger signal": when startled or frightened, a swimming beaver will rapidly dive while forcefully slapping the water with its broad tail. This creates a loud "slap", audible over large distances above and below water. This noise serves as a warning to beavers in the area. Once a beaver has made this danger signal, nearby beavers dive and may not reemerge for some time. Beavers are slow on land, but good swimmers that can stay under water for as long as 15 minutes. (Wilson, 1971) Rarely, a frightened beaver attacks a human.

Beavers do not hybernate, but store sticks and logs underwater to feed on during the winter.

Fossil remains of beavers are found in the peat and other superficial deposits of England and the continent of Europe; while in the Pleistocene formations of England and Siberia occur remains of a giant extinct beaver, Trogontherium cuvieri, representing a genus by itself.

Beavers have webbed hind-feet, and a broad, scaly tail. They have poor eyesight, but keen senses of hearing, smell, and touch.

Beavers continue to grow throughout life. Adult specimens weighing over are not uncommon. Females are as large as or larger than males of the same age, which is uncommon among mammals.

Etymology

The word is descended from the Proto-Indo-European name of the animal, cf. Sanskrit babhru's, brown, the great ichneumon, Lat. fiber, Ger. Biber, Swed. bäver, Russ. bobr'; the root bhru has given "brown," and, through Romanic, "bronze" and "burnish."

Species

They are the only extant members of the family Castoridae, which contains a single genus, Castor. Genetic research has shown the European and North American beaver populations to be distinct species and that hybridization is unlikely. Beavers are closely related to squirrels (Sciuridae), agreeing in certain structural peculiarities of the lower jaw and skull. In the Sciuridae the two main bones (tibia and fibula) of the lower half of the leg are quite separate, the tail is round and hairy, and the habitats are arboreal and terrestrial. In the beavers or Castoridae these bones are in close contact at their lower ends, the tail is depressed, expanded and scaly, and their habitats are aquatic.

European and American beavers grow to about long (plus of tail). They are essentially aquatic in their habits, never traveling by land unless driven by necessity. They are crepuscular rather than nocturnal, which means they are active at dawn and dusk, and subsist chiefly on bark and twigs or the roots of water plants. They have been known to eat grasses on the banks of rivers and streams.

European Beaver

The European Beaver (Castor fiber) was hunted almost to extinction in Europe, both for fur and for castoreum, a secretion of its scent gland believed to have medicinal properties. However, the beaver is now being re-introduced throughout Europe. Several thousand live on the Elbe, the Rhone and in parts of Scandinavia. A thriving community lives in northeast Poland, and the European Beaver also returned to the Morava River banks in Slovakia and the Czech Republic. They have been reintroduced in Scotland, Bavaria, Austria, The Netherlands and Serbia (Zasavica bog) and are spreading to new locations.

The beaver became extinct in Great Britain in the sixteenth century: Giraldus Cambrensis reported in 1188 (Itinerarium ii.iii) that it was to be found only in the Teifi in Wales and in one river in Scotland, though his observations are clearly first hand. In October 2005, six European beavers were reintroduced to Britain in Lower Mill Estate in Gloucestershire; in July 2007 a colony of four European beavers was established at Martin Mere in Lancashire, and there are plans for re-introductions in Scotland and Wales.

American Beaver

The American Beaver (Castor canadensis), also called the Canadian Beaver (which is also the name of a subspecies), or simply Beaver in North America, is native to Canada, much of the United States and parts of northern Mexico. The chief feature distinguishing C. canadensis from C. fiber is the form of the nasal bones of the skull. This species was introduced to the Argentine and Chilean Tierra del Fuego, as well as Finland, France, Poland and Russia.

The American beaver's preferred food is the water-lily (Nuphar luteum), which bears a resemblance to a cabbage-stalk, and grows at the bottom of lakes and rivers. Beavers also gnaw the bark of birch, poplar, and willow trees; but during the summer a more varied herbage, with the addition of berries, is consumed.

These animals are often trapped for their fur. During the early 19th century, trapping eliminated this animal from large portions of its original range. However, through trap and transfer and habitat conservation it made a nearly complete recovery by the 1940s. Beaver furs were used to make clothing and top-hats. Much of the early exploration of North America was driven by the quest for this animal's fur. Native peoples and early settlers also ate this animal's meat. The current beaver population has been estimated to be 10 to 15 million; one estimate claims that there may at one time have been as many as 90 million.

Giant beaver

The North American Giant beaver (Castoroides ohioensis) was one of the largest rodents that ever evolved. About the size of a Grizzly Bear, it disappeared along with other large mammals in the Holocene extinction event, which began about 13,000 years ago.

Habitat

The habitat of the beaver is the riparian zone, inclusive of stream bed. The actions of beavers for hundreds of thousands of years in the Northern Hemisphere have kept these watery systems healthy and in good repair, although a human observing all the downed trees might think that the beavers were doing just the opposite.

The beaver works as a keystone species in an ecosystem by creating wetlands that are used by many other species. Next to humans, no other extant animal appears to do more to shape its landscape.

Effect of beaver on trees

Beavers fell trees for several reasons. They fell large mature trees, usually in strategic locations, to form the basis of a dam. They fell small trees, especially young second growth trees, for food. Ponds created by beavers also kill trees by drowning.

The dam's primary role: the beaver's home

Beaver dams are created as a protection against predators, such as coyotes, wolves and bears, and to provide easy access to food during winter. Beavers always work at night and are prolific builders, carrying mud and stones with their fore-paws and timber between their teeth. Because of this, destroying a beaver dam without removing the beavers is difficult, especially if the dam is downstream of an active lodge. Beavers can rebuild such primary dams overnight, though they may not defend secondary dams as vigorously. (Beavers may create a series of dams along a river.)

Shape of dam

A beaver shapes a dam according to the strength of the water's current. Relatively still water encourages dams that are almost straight; while dams in stronger currents are curved, bowed toward upstream. The beavers use driftwood, green willows, birch and poplars; and they mix in mud and stones that contribute to the dam's strength.

Beavers are known to build very large dams. The largest known was discovered by satellite imagery in Northern Alberta in 2007, approximately long , beating the previous record holder found near Three Forks, Montana, at long, high, and thick at the base. When objectionable beaver flooding occurs, modern water level control devices can be installed for a cost-effective and environmentally sound solution. Unwanted damage to trees can be prevented by wrapping chicken wire or sheet metal around the base of trees.

"In places," writes Hearne, "which have been long frequented by beavers undisturbed, their dams, by frequent repairing, become a solid bank, capable of resisting a great force both of ice and water; and as the willow, poplar and birch generally take root and shoot up, they by degrees form a kind of regular planted hedge, which I have seen in some places so tall that birds have built their nests among the branches."

Stimulus for dam-building

It is primarily prolonged exposure to the sound of water in motion that stimulates the beavers to build. However, studies involving beaver habitual activities have indicated that beavers may respond to an array of stimuli, not just the sound of running water (such as seeing water movement). In two experiments Wilson (1971) and Richard (1967, 1980) demonstrate that, although beavers will pile material close to a loudspeaker emitting sounds of water running, they only do so after a considerable period of time. Additionally the beavers, when faced with a pipe allowing water to pass through their dam, eventually stopped the flow of water by plugging the pipe with mud and sticks. The beavers were observed to do this even when the pipe extended several meters upstream and near the bottom of the stream and thus produced no sound of running water. Beavers normally repair damage to the dam and build it higher as long as the sound continues. However, in times of high water, they often allow spillways in the dam to flow freely.

Disruption by dams

Beaver dams can be disruptive; the flooding can cause extensive property damage, and when the flooding occurs next to a railroad roadbed, it can cause derailments by washing-out the tracks, or when a beaver dam bursts and the resulting flash flood overwhelms a culvert. This disruption is not limited to human geography; beavers can destroy nesting habitat for endangered species, and often destroy mature trees for which they have no use. Introduced to an area without its natural predators, as in Tierra del Fuego, beavers have flooded thousands of acres of land and are considered a plague. One notable difference in Tierra del Fuego from most of North America is that the trees in Tierra del Fuego do not coppice as do willows, poplars, aspens, and other North American trees. Thus the "damage" by the beavers seems more severe.

Benefits of dams compared to disruption

On the other hand, dam building can be beneficial in restoring wetlands. Such wetland benefits include flood control downstream, biodiversity (by providing habitat for many rare as well as common species), and water cleansing, both by the breakdown of toxins such as pesticides and the retention of silt by beaver dams. Over the eons, this collection of silt produces the rich bottom land so sought after by farmers. Beaver dams reduce erosion as well as decrease the turbidity that is a limiting factor for much aquatic life. While beavers can create damage, part of the problem is one of perception and time scale. Such damage as the undermining of a roadway or the drowning of some trees is very visible shortly after the beginning of beavers activity in an area. The benefits may be long-term and unnoticed except, for example, by someone monitoring a catchment.

The dam's role in the stream's life cycle

Wetland creation

If a beaver pond becomes too shallow due to sediment accumulation, or if the tree supply is depleted, beavers will abandon the site. Eventually the dam will be breached and the water will drain out. The rich thick layer of silt, branches, and dead leaves behind the old dam is the ideal habitat for wetland species. Many may have been on the fringes of the pond. Wetlands have significant environmental benefits.

The grazing meadow (vega and valle)

As the wetland fills and dries out, pasture species colonize it and it becomes a meadow suitable for grazing. In an area with nothing but forest down to the stream edge, this provides a valuable niche for many animals which otherwise would be excluded.

The riverine forest

Finally the meadow will be colonized by riverine trees, typically aspens, willows and such species which are favoured by the beaver. Beavers are then likely to recolonize the area, and the cycle begins again.

The dam's beneficial ecological effects

Bottom land

Each time the stream life cycle repeats itself another layer of rich organic soil is added to the bottom of the valley. The valley slowly fills and the flat area at the bottom gets wider. Research is sparse, but it seems likely that much of the fabled bottom land in North America was created, or at least added to, by the efforts of the generations of beavers that lived there.

Flood control

A beaver dam has a certain amount of freeboard above the water level. When heavy rains occur, the pond fills up and the dam gradually releases the extra stored water. Often this is all that is necessary to reduce the height of the flood wave moving down the river, and will reduce or eliminate damage to human structures. Flood control is achieved in other ways as well. The surface of any stream intersects the surrounding water table. By raising the stream level, the gradient of the surface of the water table above the beaver dam is reduced, and water near the beaver dam flows more slowly into the stream. This further helps in reducing flood waves, and increases water flow when there is no rain. Beaver dams also smooth out water flow by increasing the area wetted by the stream. This allows more water to seep into the ground where its flow is slowed. This water eventually finds its way back to the stream. Rivers with beaver dams in their head waters have lower high water and higher low water levels.

Nutrient removal

Beaver ponds can cause the removal of nutrients from the stream flow. Farming along the banks of rivers often increases the loads of phosphates, nitrates and other nutrients, causing problems downstream when this water is extracted for drinking. Besides silt, the beaver dam collects twigs and branches from the beavers' activity and leaves, notably in the fall. The main component of this material is cellulose, a polymer of β-glucose monomers (This creates a more crystalline structure than is found in starch, which is composed of α-glucose monomers. Cellulose is a type of polysaccharide.) Many bacteria produce cellulase which can split off the glucose and use it for energy. Just as algae get their energy from sunlight, these bacteria get their energy from cellulose, and they form the base of a very similar food chain. However, a source of energy is not enough for growth. These bacterial populations face serious shortages of nitrous and phosphorous compounds, and will absorb these nutrients as they pass by in the water stream. In this way, these and other nutrients are fixed into the beaver pond and the surrounding ecology, and are removed from the stream.

Pesticide and herbicide removal

Agriculture also introduces herbicides and pesticides into our streams. Bacteria are an extremely variable lot and some of these toxicants are metabolized and decomposed by the bacteria in the cellulose-rich bottom of a beaver dam.

Denitrification

Some scientists believe that the nitrate cascade, the production of far more fixed nitrogen than the natural cycles can turn back into nitrogen gas, may be as much of a problem to our ecology as carbon dioxide production. It is likely, but not proven, that beaver dams along a stream may contribute to denitrification (the conversion of nitrogen compounds back into nitrogen). In sewage plants, denitrification is achieved by passing the water through successive aerobic and anaerobic stages. Under a beaver dam, as the water seeps down into the soil, the oxygen is consumed by the fauna in the rich organic layer. At some point all the oxygen is used up and the soil becomes anaerobic. This water eventually finds its way back into the aerobic stream and into another beaver dam. This aerobic, anaerobic cycle continues all the way down the stream and denitrification is a likely result.

Beavers and salmon

Beaver dams are a nursery for salmon. An early indication of this was seen following the 1818 agreement between the British government of Canada and the government of America allowing Americans access to the Columbia watershed. The Hudson's Bay Company, in a fit of pique sent word to its trappers to extirpate the fur-bearing animals in the area. The beaver was the first to go. Salmon runs fell precipitously in the following years even though, at that time, none of the factors were extant that we associate with the decline of salmon runs.

The functions of beaver dams in increasing salmon runs are many. They provide deep enough water for the juvenile salmon to hide from predatory wading birds. They trap nutrients in their ecology and notably the huge nutrient pulse represented by the migration of the adult salmon upstream. These nutrients help feed the juveniles after they finish their yolk. They provide quiet water so that the young salmon can put energy into growth rather than into fighting currents and larger smolt with a food reserve have a better chance when they reach the sea. And beaver dams keep the water clear which favours all the salmonoids, trout included.

Lodges

The ponds created by well-maintained dams help isolate the beavers' home, their lodge, which is also created from severed branches and mud. The beavers cover their lodges late every autumn with fresh mud which freezes when the frost sets in. The mud becomes almost as hard as stone, so that neither wolves nor wolverines can get in.

The lodge has underwater entrances to make entry nearly impossible for any other animal (however, muskrats have been seen living inside beaver lodges with the beavers who made it). A very small amount of the lodge is actually used as a living area. Contrary to popular belief, beavers actually dig out their den with an underwater entrance after they finish building the dam and lodge structure. There are typically two dens within the lodge, one for drying off after exiting the water, and another, drier one where the family actually lives.

Their houses are formed of the same materials as the dams, with little order or regularity of structure, and seldom contain more than four old, and six or eight young beavers. Sometimes some of the larger houses have one or more partitions, but these are only posts of the main building left by the builders to support the roof, for the apartments have usually no communication with each other except by water.

When the ice breaks up in spring they always leave their embankments, and rove about until a little before fall, when they return to their old habitations, and lay in their winter stock of wood. They seldom begin to repair the houses till the frost sets in, and never finish the outer coating till the cold becomes severe. When they erect a new habitation they fell the wood early in summer, but seldom begin building till towards the end of August.

Invasiveness

Beavers can create serious damage when spread outside of their natural environment, and are therefore considered in some places as pests or invasive species. For example, in the 1940s, beavers were brought to the island of Tierra Del Fuego in southern Argentina, for commercial fur production. However, the project failed and the beavers were released into the wild. Having no natural predators in their new environment, they quickly spread throughout the island, reaching a number of 100,000 individuals within just 50 years (when released into the wild there were only a few pairs). They are now considered a serious invasive species on the island, due to their massive destruction of forest trees, and efforts are being made for their eradication.

Commercial uses

Beaver pelts were used for barter by Native Americans in the 17th century to gain European goods. They were then shipped back to Great Britain and France where they were made into clothing items. Widespread hunting and trapping of beavers led to their endangerment. Eventually, the fur trade declined due to declining demand in Europe and the takeover of trapping grounds to support the growing agriculture sector. A small resurgence in beaver trapping has occurred in some areas where there is an over-population of beaver; trapping is done when the fur is of value, the remainder of the animal may be used as feed. In the 1976/1977 season, 500,000 beaver pelts were harvested in North America.

Both beaver testicles and castoreum, a bitter-tasting secretion with a slightly fetid odor contained in dried preputial or vaginal follicles of male or female beaver, have been articles of trade for use in traditional medicine. Yupik (Eskimo) medicine used dried beaver testicles like willow bark to relieve pain. Beaver testicles were exported from Levant (a region centered on Israel) from the tenth to nineteenth century. Claudius Aelianus comically described beavers chewing off their testicles to preserve themselves from hunters. European beavers (Castor fiber) were eventually hunted nearly to extinction in part for the production of castoreum, which as used as an analgesic, anti-inflammatory, and antipyretic. Castoreum was described in the 1911 British Pharmaceutical Codex for use in dysmenorrhea and hysterical conditions (i.e. pertaining to the womb), for raising blood pressure and increasing cardiac output. The activity of castoreum has been credited to the accumulation of salicin from willow trees in the beaver's diet, which is transformed to salicylic acid and has an action very similar to aspirin. Castoreum continues to be used in perfume production.

In culture

As a national emblem

The importance of the American Beaver in the development of Canada through the fur trade led to its designation as the national animal. It is depicted on the Canadian five-cent piece and was on the first postage stamp issued in the Canadian colonies in 1849 (the so-called "Three-Penny Beaver"). As a national symbol, the beaver was chosen to be the mascot of the 1976 Summer Olympics held in Montreal with the name "Amik" ("beaver" in Anishinaabemowin). The beaver is also the symbol of many units and organizations within the Canadian Forces, such as on the cap badges of the Royal 22^e Régiment and the Canadian Military Engineers.

In dietary law

In the 17th century, based on a question raised by the Bishop of Quebec, the Roman Catholic Church ruled that the beaver was a fish (beaver flesh was a part of the indigenous peoples' diet, prior to the Europeans' arrival) for purposes of dietary law. Therefore, the general prohibition on the consumption of meat on Fridays during Lent does not apply to beaver meat. The legal basis for the decision probably rests with the Summa Theologica of Thomas Aquinas, which bases animal classification as much on habit as anatomy.

References

• ITIS 180211 2002-12-14
• The American Beaver and His Works by Lewis Henry Morgan, Published by J.B. Lippincott & Co., 1868. Some 1911 Britannica material appears to be copied from this source.

Further reading

• Rue, Leonard Lee, III.
• The World of the Beaver, Lippincott Company, 1964.
• Beavers, 2002. ISBN 0896585484
• The Tent Dwellers by Albert Bigelow Paine. Beavers' habits, habitat and conservation status (as of 1908) are recurring themes.

External links

• Beaver Facts & Pictures
• Beavers: Wetlands & Wildlife website
• The Romance of the Beaver - history of the beaver in the western hemisphere by A. Radclyffe Dugmore.
• Aigas Field Centre Beaver Project - history of a pair of European beavers released into a large enclosure in the Highlands of Scotland.