No. 4 Flying Training School takes RAF and Royal Navy pilots from 1FTS at RAF Linton-on-Ouse and trains them to fly fast jets, prior to training on an Operational Conversion Unit. 4 FTS is divided into two squadrons; 208 Sqn provides the advanced flying training, students then moving onto 19 Sqn to receive tactics and weapons training.
Valley is also home to 'C' Flight of 22 Sqn with Sea King helicopters. These are busy in the Search and Rescue role, rescuing people from ships in the Irish Sea, from the mountains of nearby Snowdonia and elsewhere. The mountain rescue work in Snowdonia is coordinated with the Wales Mountain Rescue Association
The base is also home to SARTU (search and rescue training unit) using Squirrel and Griffin helicopters, and newly home to the relocated headquarters elements of both 22 and 202 Squadrons.
RAF Mona, also on Anglesey, acts as a relief landing ground.
A terminal facility, costing £1,500,000, has been constructed at RAF Valley, which will be named Anglesey Airport (Maes Awyr Môn); the capacity of which is projected to handle up to ten such PSO flights, each day. Inverness based Highland Airways operates the Cardiff route, using BAe Jetstream 31 aircraft. The first public flight took place on 8 May 2007.