pulse, alternate expansion and contraction of artery walls as heart action varies blood volume within the arteries. Artery walls are elastic. Hence they become distended by increased blood volume during systole, or contraction of the heart. During diastole, or relaxation of the heart, blood volume in the arteries decreases and the walls contract, propelling the blood farther along the arterial pathway. The effect is that of a pressure wave initiated by the heartbeat and traveling from the aorta, the major artery leaving the heart, along the walls of all the other arteries. It takes about a quarter of a second for this wave to travel from the aorta to the arteries in the soles of the feet. The rate of heartbeat is equivalent to the pulse rate. Usually the pulse rate is determined by counting the pulsations per minute in the radial artery at the wrist. It may also be determined at any other artery point near the surface of the body. The normal rate is 70 to 90 pulsations per minute in adults, and 90 to 120 in children. Various diseases may be indicated by changes in the rate, rhythm, and force of the pulse.
pulse, in botany, common name for members of the Fabaceae (Leguminosae), a large plant family, called also the pea, or legume, family. Numbering about 650 genera and 17,000 species, the family is third largest, after the asters and the orchids. Some botanists divide the Fabaceae into three or more separate families, but most species share certain common and easily recognizable features. The leaves are usually compound; the fruit is a legume (a type of pod); and the blossoms may have an irregular butterflylike (papilionaceous) shape. Typically, the flowers have 10 stamens, and the corolla and the calyx are formed of 5 petals and 5 sepals, respectively. Some species have thorny branches.

The Fabaceae include herbs, shrubs, and trees distributed throughout the world in a great variety of forms. Arboreal species occur in temperate and, frequently, in tropical zones, where epiphytic and climbing forms also thrive. Many leguminous shrubs and trees inhabit desert and semiarid regions, usually forming the characteristic vegetation—e.g., the acacias of the S African bushveld and of Australia, and the mesquite of the American Southwest.

The Pulses and Their Uses

Economically, the family is second only to the grasses in importance. Legumes provide valuable and nutritive foods because the food stored for the embryo in the seed (e.g., the pea) is rich in protein. In many regions, especially where meat is scarce or expensive, legumes—notably peas, beans, lentils, peanuts, carob, and soybeans—are staples of the diet. The Fabaceae are equally important as fodder and forage plants; clover, alfalfa, vetch, lupine, beggarweed, lespedeza, sainfoin, and soybeans are among the numerous valuable types.

These food and forage legumes are chief among the plants used as "green manure" (see manure). Nitrogen-fixing bacteria dwelling in nodules of the roots of most legumes fix free nitrogen from the air into the nitrogenous compounds needed by all forms of life for building proteins (see nitrogen cycle). Rotation of leguminous crops with nonleguminous crops has long been a standard agricultural practice; the soil is enriched when their roots are left to decay after harvesting.

The pulse family also provides gums and resins (e.g., tragacanth, copal, and acacia and carob gums), dyes and tannins (e.g., from the indigo plant, logwood, brazilwood, and types of acacia and broom), timber (e.g., rosewood, locust, honey locust, and acacia), medicines (e.g., from tamarind, licorice, and senna), perfume oils (e.g., from acacia, black locust, broom, and sweet pea), vegetable oils (e.g., soybean and peanut oils), and other commercial items such as flavorings, fibers, and insecticides.

In many parts of the world native species of the Leguminosae are of great importance locally, if not commercially. Often every part of the plant finds some use: the pods and leaves for food, beverages, and forage; the wood and stems for building purposes, fiber, and household items; and the leaves, blossoms, and bark for domestic remedies. The blossoms of many of the Leguminosae are excellent honey sources. Species that grow in arid climates are particularly valuable because of the scarcity of other fodder, food, and timber crops; they are also important to wildlife for forage and cover. Native Americans have cultivated bean plants since antiquity and still rely on breadroot, redwood, mesquite, and many other species for food and other products.

Among the native North American trees cultivated for shade or for their beautiful springtime blossoms are the locusts, the honey locust, the yellowwood, the redbud, and the acacias. The mimosas, sennas, laburnums, poincianas, Old World acacias, shrubby brooms, and wisteria have been introduced for the same purpose. The American lupines, the Old World sweet pea, and numerous types of clover are among the cultivated herbaceous species. In all, members of over 140 genera of the Leguminosae are grown for ornament. Furze from Europe and the kudzu vine from Asia have been introduced for erosion control (the latter has become a noxious weed). The locoweeds and lupines of the western states are among the plants poisonous to livestock.

See articles on individual plants.


The pulse family is classified in the division Magnoliophyta, class Magnoliopsida, order Rosales.

In medicine, a person's pulse is the throbbing of their arteries. It can be felt in any place that allows for an artery to be compressed against a bone, such as at the neck (carotid artery), at the wrist (radial artery), behind the knee (popliteal artery), on the inside of the elbow (brachial artery), and near the ankle joint (posterior tibial artery). The pulse rate can also be measured by measuring the heart beats directly (the apical pulse).

Pressure waves move the artery walls, which are pliable; these waves are not caused by the forward movement of the blood. When the heart contracts, blood is ejected into the aorta and the aorta stretches. At this point, the wave of distention (pulse wave) is pronounced but relatively slow-moving (3–6 m/s). As it travels towards the peripheral blood vessels, it gradually diminishes and becomes faster. In the large arterial branches, its velocity is 7–10 m/s; in the small arteries, it is 15–35 m/s. The pressure pulse is transmitted fifteen or more times more rapidly than the blood flow.

Pulse is also used, although incorrectly, to denote the frequency of the heart beat, usually measured in beats per minute. In most people, the pulse is an accurate measure of heart rate. Under certain circumstances, including arrhythmias, some of the heart beats are ineffective, and the aorta is not stretched enough to create a palpable pressure wave. The pulse is too irregular and the heart rate can be (much) higher than the pulse rate. In this case, the heart rate should be determined by auscultation of the heart apex, in which case it is not the pulse. The pulse deficit (difference between heart beats and pulsations at the periphery) should be determined by simultaneous palpation at the radial artery and auscultation at the heart apex.

A normal pulse rate for a healthy adult, while resting, can range from 60 to 100 beats per minute (BPM), although well-conditioned athletes may have a healthy pulse rate lower than 60 BPM. Bradycardia occurs when the pulse rate is below 60 per minute, whereas tachycardia occurs when the rate is above 100 BPM. During sleep, the pulse can drop to as low as 40 BPM; during strenuous exercise, it can rise as high as 150–200 BPM. Generally, pulse rates are higher in infants and young children. The resting heart rate for an infant is usually close to an adult's pulse rate during strenuous exercise (average 110 BPM for an infant).

A collapsing pulse is a sign of hyperdynamic circulation.

Common pulse sites

  • Axillary pulse: located inferiorly of the lateral wall of the axilla
  • Apical pulse: located in the 4th or 5th left intercostal space, just to the left of the sternum. In contrast with other pulse sites, the apical pulse site is unilateral, and measured not over an artery, but over the heart itself (more specifically, the apex of the heart).
  • Brachial pulse: located between the biceps and triceps, on the medial side of the elbow cavity, frequently used in place of carotid pulse in infants (brachial artery)
  • Carotid pulse: located in the neck (carotid artery). The carotid artery should be palpated gently and while the patient is sitting or lying down. Stimulating its baroreceptors with vigorous palpitation can provoke severe bradycardia or even stop the heart in some sensitive persons. Also, a person's two carotid arteries should not be palpated at the same time. Doing so may limit the flow of blood to the head, possibly leading to fainting or brain ischemia. It can be felt between the anterior border of the sternocleidomastoid muscle, above the hyoid bone and lateral to the thyroid cartilage.
  • Dorsalis pedis pulse: located on top of the foot (dorsalis pedis artery).
  • Facial pulse: located on the mandible (lower jawbone) on a line with the corners of the mouth (facial artery).
  • Femoral pulse: located in the thigh, halfway between the pubic symphysis and anterior superior iliac spine (femoral artery).
  • Popliteal pulse: located behind the knee in the popliteal fossa, found by holding the bent knee. The patient bends the knee at approximately 120°, and the physician holds it in both hands to find the popliteal artery in the pit behind the knee.
  • Radial pulse: located on the thumb side of the wrist (radial artery). It can also be found in the anatomical snuff box.
  • Temporal pulse: located on the temple directly in front of the ear (superficial temporal artery).
  • Tibialis posterior pulse: located on the medial side of the ankle (facing inwards) behind the medial malleolus (posterior tibial artery).
  • Ulnar pulse: located on the little finger side of the wrist(ulnar artery).

The ease of palpability of a pulse is dictated by the patient's blood pressure. If his or her systolic blood pressure is below 80 mmHg, the radial pulse will not be palpable. Below 70 mmHg, the brachial pulse will not be palpable. Below 60 mmHg, the carotid pulse will not be palpable. However, a study by the National Institutes of Health indicated that this method was not accurate enough and often overestimated a patient's systolic blood pressure. The lack of a palpable carotid pulse is often an indication of death.

Reading a pulse

Pulses are manually palpated with fingers. When palpating the carotid artery, the femoral artery or the brachial artery, the thumb may be used. However, the thumb has its own pulse which can interfere with detecting the patient's pulse at other points, where two or three fingers should be used. Fingers or the thumb must be placed near an artery and pressed gently against a firm structure, usually a bone, in order to feel the pulse.

Make sure the person is calm and has been resting for 5 minutes before reading the pulse. Put the index and middle fingers over the pulse count, and count for 30 seconds, and afterwards multiply by 2, to get the pulse rate. If the person's pulse rate is irregular, count for a full minute, and do not multiply. Averaging multiple readings may give a more representative figure.

Home blood pressure measurement devices also typically give a pulse reading.


  • Jarvis, Caroylin (2003). Physical Examination & Health Assessment. Saunders.

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