Dinosaurs in the game take the place of traditional wildlife and livestock, though the dinosaurs in-game are highly stylized and not necessarily scientifically accurate. Wandering dinosaurs can be hunted for food, though some are dangerous and aggressive species will attack any player unit that wanders too close. Other prehistoric creatures make an appearance as well, for example woolly mammoths are used where other games might feature elephants.
The game borrows the general counter system from AoE: regular infantry are countered by mounted units, mounted units are countered by pikemen, and pikemen are countered by regular infantry. Heroes and advanced units are more powerful overall.
Players advance through epochs in order to unlock new technologies and upgrades.
Later, three young scientists are led to Jarvis Babbit, a mysterious man who seems to know something about a series of oddities that the scientists have come across in their research. The American geologist Anthony Cole, Swedish biologist Stina Holmlund, and Hungarian physics genius Béla Andràs Benedek are sent there, seemingly for an expedition. The 19th-century scientists find out about this, and try to prevent the three scientists from releasing information about this parallel dimension by trapping them in the hostile ParaWorld.
The three scientists serve as the game's core "hero" units. Their main goal is to return to their home universe, but to do this they will need to enlist the aid of enlightened natives and battle against the hostile creatures, tribes, and ultimately the masterminds trying to stop them. Along the way they'll meet several characters who are seemingly based on classical scientists such as Nicola Tesla ("Nikolaj Taslow" in the game). They eventually discover that a man named David Leighton, who is one of Babbit's men, is orchestrating events on the parallel world, and pursue him. They also learn of a plot to permanently seal the world away by moving the planet out of orbit, which will also trigger a cataclysm.
In order to save the planet and secure their own escape from ParaWorld, they are forced to assault Babbit's fortifications. They manage to stop the countdown and Stina cuts off Leighton's arms. Babbit makes a last-ditch effort to stop the heroes by using a giant robot. Babbit falls into lava and presumably dies. The heroes finally go back home. However, Cole is awakened in the middle of the night by a phone call telling him that the dinosaurs have come through as well. The chaos of ParaWorld now ensues on planet Earth.