|First Appearance:||The Wood In Danger (1x01)|
|Last Appearance:||New Enemies (2x04)|
|Offspring:||Mossy, and an unnamed female|
He often believes that he is a burden, going so far as to slip off Badger's back and try walking on his own. Despite this, he does prove useful to the group on several occasions; once using his digging ability to free them from imprisonment in a barn. In the town, he finds a hole in the church wall, allowing the animals to enter and take shelter from a storm.
Mole is very greedy, particularly when it comes to worms, and this greed sometimes gets him into trouble; once causing him to be lost in a forest fire. As a result of this, Mole is rescued by a fireman and then escapes, to find himself unable to see the other animals. Owl and Kestrel then put on a flying display to distract the human firefighters.
Their friendship is so strong that Badger is not told of mole's death, for fear that the pain will be too much for him.
Mole also finds a mate, Mateless, in White Deer Park. He meets her during the winter, asking her to join him in his burrow to keep warm. As winter comes to an end Mole is seen dancing with Mateless under a winter moon, until a pile of melting snow falls on them. Both also appear celebrating when Fox triumphs over the poachers, although this may be a mistake because the animals later do not know Mole had found a mate.
With his mate, Mole fathers a son, Mossy, and an unnamed daughter.