Loblolly-bay (Gordonia lasianthus), also called holly-bay, gordonia, and bay, is a small to medium-sized evergreen tree or shrub found in acid, swampy soils of pinelands and bays on the Atlantic and Gulf Coastal Plains. It is slow growing with soft, light-colored, fine-grained wood of little commercial value, although loblolly-bay could be managed as a source of pulpwood. The white showy flowers and shiny foliage make it a desirable ornamental, but it is not easy to cultivate. Deer browse stump sprouts heavily.
|Temperature:||16 °C (60 °F)||21 °C (70 °F)|
|Annual daily average||3 °C (37 °F)||11 °C (52 °F)|
|Normal daily minimum; January||31 °C (87 °F)||32 °C (90 °F)|
|Normal daily maximum; July||11 °C (52 °F)||10 °C (50 °F)|
|Days with daily max. above
32 °C (90 °F)
|Days with daily min. below
0 °C (32 °F)
|Mean frost-free days||230||320|
Approximately 53 percent of the annual precipitation occurs during the months of June, July, August, September, and October. Annually there are from 110 to 120 days with only a trace of rainfall.
Loblolly-bay grows in flat woodlands or shallow depressions with little or no slope, slow runoff, rapid permeability, and poor to very poor drainage. In South Carolina the soils are usually of sandy coastal plain or marine origin, except for the organic soils. The water table is at or near the soil surface for 6 to 9 months of the year.
In South Carolina's lower Coastal Plain, loblolly-bay is found in wet flats and in bays, typically the Carolina Bays (11). In the upper and middle Coastal Plain, it is found mainly along the edges of Carolina Bays and is widely dispersed in wet, flat woodlands on certain soil types.
Loblolly-bay is a minor component of Loblolly Pine-Hardwood (Type 82) but cannot be found consistently. In the middle Coastal Plain of South Carolina, loblolly-bay is found with loblolly pine, water oak (Quercus nigra), sweetgum (Liquidambar styraciflua), American holly (Ilex opaca]]), redbay, longleaf pine (Pinus palustris), and yellow-poplar (Liriodendron tulipifera). Loblolly-bay is found only in the wetter areas of this forest cover type.
Pond Pine (Type 98) is the typical vegetation of wet flats and some Carolina Bays in South Carolina. Loblolly-bay, pond pine, sweetbay, and redbay are the tree species present, and they rarely form a closed canopy. The thick, shrub layer is composed of fetterbush, greenbrier vines, inkberry, and loblolly-bay.
Sweetbay-Swamp Tupelo-Redbay (Type 104) is the "broadleaf evergreen forest" of the lower Coastal Plain of North and South Carolina. Loblolly-bay is a minor component in the overstory along with red maple (Acer rubrum), black tupelo (Nyssa sylvatica var. sylvatica), sweetgum, and water oak.
Flowering and fruiting- Flowers are perfect. Flower bud formation is visible by the time new leaves fully expand. The peduncle expands rapidly and the young bud slowly enlarges until it opens. Flower buds at the top of the tree open first. Flowers are first seen from the last week in June to the first week of July and may be seen until mid-August. Flowers remain open for 1 or 2 days and are pollinated by humble bees, thrips, flies, and hummingbirds. After the second day the sepals and petals fall, leaving the ovary at the end of the peduncle.
Seed production and dissemination- As the ovaries develop they gradually turn brown and five sutures develop. Mature, open capsules are first seen during September or October, and all of the capsules open by the middle of December. Seeds are shaken out of the capsules by the wind and empty capsules remain attached until peduncle and capsule abscission, which first occurs about the last of December and continues through the winter.
Seedfall starts in October, peaks in December, and continues until the first of March. Loblolly-bay seeds are light (120,000 to 151,000/lb or 265,000 to 333,000/kg) and winged. Results from one study indicated that approximately 99 percent of the seeds produced fall within two tree heights of the source tree. This study also indicated that this distribution of seedfall is concentrated near the source tree, 60 percent of the seeds falling within a distance equal to one-half of the tree height, and 94 percent of the seeds falling within a distance equal to the tree height. Seedfall rates during a 2-year study varied from 2,600 to 273,000/ha (1,070 to 110,400/acre).
Seedling development- Seed germination in petri dishes in sunlight is high: 70 to 80 percent within 10 days. In a greenhouse heated at 13 °C to 16 °C (55 °F to 60 °F), similar germination percentages were obtained but up to 24 days were required. Germination is epigeal. Very few seedlings have been observed in the field and most of those seen apparently do not live past the first season. Loblolly-bay seedlings seem to require relatively open conditions and exposed soil for establishment. Older seedlings have only been observed where the mineral soil has been disturbed such as in recently plowed fire lines.
Initial growth of the seedlings is slow. Field observations indicated that by the end of the third growing season the seedlings were about 10 to 15 cm (4 to 6 in) tall and by the eighth growing season they were only 30 to 40 cm (12 to 16 in) high.
Vegetative reproduction- Vegetative propagation of first-year shoots in a peat and sand medium under mist is commonly used by horticulturists (1). In the field, vegetative regeneration appears to be more common than regeneration from seed. Stump sprouts may grow as much as 1 in (3 ft) the first year after the tree is cut. These stump sprouts appear to be very attractive to deer and heavy browsing has been noticed.
Numerous root-collar sprouts are produced when the trees are killed by fire (9) or if the root system is mechanically damaged by a logging or disking operation.
Rooting habit- The root system of loblolly-bay appears to reflect its strong tendency to reproduce by sprouting. A number of specimens examined had a large primary lateral root with secondary roots branching downward.
Reaction to competition- Loblolly-bay is classed as tolerant of shade. In bays and wet flats, where the tree cover is relatively light, loblolly-bay is a strong competitor. It generally increases in height faster than the pines on the adjacent upland. However, if loblolly-bay is overtopped, older trees will lose their characteristic conical shape and the crown will break up.
Damaging agents- Only two symptoms of insects or pathogens have been observed locally. Neither causal agent was identified. An ooze was noticed in a wound at the base of a mature tree, but otherwise the tree appeared healthy. An unknown grazing insect consumed all but the leaf veins of the late-season flush of leaves during August. Another noticeable sensitivity is to fire. The thin bark and shallow root system of loblolly-bay probably contribute to its low fire tolerance.
Because of its ability to grow in wet bogs and flats where loblolly pine does poorly, loblolly-bay silviculture may offer a management alternative for such areas.
Laboratory papermaking tests conducted recently and other results reported in the literature (3,4) indicate that the pulp yield from loblolly-bay was acceptable (52 percent), the bulk of the paper was low (1.46 cm/g or 2.53 in/oz), and the strength acceptable. One laboratory test indicated a breaking length of 11,525 m (37,800 ft), a tensile strength of 10.2 kgf/15 mm (38.1 lbf/in), and a count of 836 folds using the Massachusetts Institute of Technology paper folding tester (1 kg or 2.2 lb). Although further testing needs to be done, these tests do not indicate any problems in making kraft paper from loblolly-bay pulp. Pulp mills in the lower Coastal Plain of South Carolina include loblolly-bay in their hardwood pulp.
(Silvics of North America; volume 2: Hardwoods, United States Department of Agriculture Forest Service Agriculture Handbook 654, 1990).